Grizzly Bears
In a 1998 study published in the journal "Ursa," researchers studied interactions between grizzly bears and cougars in Glacier National Park, Montana, from 1992 to 1995 and in Yellowstone National Park, Wyoming, from 1990 to 1995. The study found bears often displaced cougars from their kills. It also found cougars displaced from their kills by bears lost an average of 17 to 26 percent of their daily energy requirements. Such encounters are only likely to occur when competition for food becomes intense.
Gray Wolves
In a study published in "Ecoscience," researchers detailed their studies into interactions between cougars and gray wolves in Banff National Park, Alberta, over a 12-year period from 1993 to 2004. The study concluded that wolves were responsible for cougar mortality and usurping prey carcasses from cougars, but cougars failed to exhibit reciprocal behavior. Cougars are susceptible to wolf attacks when the two species are forced to compete for food. Because the cougar is more powerful than the gray wolf, wolves are only a danger to cougars when they attack a solitary cougar as a pack.
Hunting
Cougars were once common throughout the U.S. However, prized by hunters and loathed by farmers for the impact they had on livestock, their numbers were severely depleted during the 19th century. By the early years of the 20th century, they were largely eliminated from the Midwest and Eastern states. The remaining cougar population has stabilized with responsible hunting, but while quotas are imposed in those states where it is legal to hunt them, cougars remain under threat from illegal poaching.
Environmental Threats
The greatest enemy of the cougar is environmental change caused by humans. Although "The International Union for Conservation of Nature (IUCN) Redlist" of threatened species does not categorize the cougar as an endagered species, it does acknowledge that the species is declining. In its 2008 "Puma Concolor Assessment," the IUCN concluded that cougars are threatened by habitat loss and fragmentation. The assessment also found that road kills are the principal cause of mortality in the endangered Florida cougar subpopulation, and heavily traveled roads are a major barrier to puma movements and dispersal.